| mTavari gverdi |
| saqarTvelos flora da fauna |
| vsaqarTvelos buneba |
| cxovelTa samyaro |
| WianWvelebi |
| WianWvelebi saqarTveloSi |
Camponotus |
1 species live in Georgia:
Hita Garcia, Wiesel and Fischer (2013) - Camponotus is an extremely large and complex, globally distributed genus. At present, more than 1000 species and nearly 500 subspecies belonging to 45 subgenera are described (Bolton, 2012) and it could well be the largest ant genus of all. The enormous species richness, high levels of intraspecific and geographic variation and polymorphism render the taxonomy of Camponotus one of the most complex and difficult. Revisionary studies on Camponotus are generally confined to species groups and/or small geographical regions (e.g. Robertson & Zachariades, 1997; Snelling, 2006; McArthur, 2007; Shattuck & Janda, 2009). These ants live in a variety of habitats and microhabitats and the sheer size of the genus makes any characterisation of their biology challenging. Nests are built in the ground, in rotten branches or twigs, or rarely into living wood (Bolton, 1973a) and most species possess a highly generalistic diet. Recently, Blaimer et al. (2015) found that Camponotus is paraphyletic with respect to Phasmomyrmex and Forelophilus, and that Colobopsis is sister to the remainder of the Camponotini while Dinomyrmex is nested outside of Camponotus. For this reason, Ward et al. (2016) revised the generic delimitation of the Camponotini (see the individual generic pages for additional information). Identification Camponotus is a hyperdiverse genus (> 1,000 species, > 400 subspecies) that is rife with taxonomic problems and puzzles. Various subgenera, complexes, and informal groupings have either been defined in taxonomic publications or have been informally recognized. All of this sorting of species at the level between the genus and species level is an attempt to introduce some order to the genus and/or allow for a more workable framework for understanding Camponotus diversity within a particular region. The Camponotus species groups page provides links to some of the Antwiki pages that include Camponotus species groupings and information. See, particularly, Emery's (1925) keys to the subgenera of Camponotus, as well as related pages linked therein. Distinguishing Colobopsis from Camponotus To differentiate Colobopsis from Camponotus Ward et al. (2016) proposed the following global key: 1 Not occurring in Fiji or New Caledonia ... 2 - Occurring in Fiji ... 3 - Occurring New Caledonia ... 4 2 Generally small species, HW 0.65–1.10 (except cylindrica-group of Southeast Asia with HW 1.20–1.70, and facies as in Figures 4 and 5); either antennal insertions relatively well separated, such that ASM/HW 0.36–0.47 and ASM/CLW 0.66–0.98, and/or clypeus relatively narrow, such that CLW/CLL 0.96–1.32; antennal insertions occurring at about midlength of frontal carinae; anterolateral extremities of clypeus set off from rest of clypeus by a sulcus or impression, so clypeus appears to lack prominent anterolateral extensions (Figures 2–5) ... Colobopsis - Small to large species, HW 0.70–3.00; antennal insertions less well separated, such that ASM/HW 0.22–0.35 and ASM/CLW 0.35–0.68; clypeus variable in shape but in smaller species with HW 0.70–1.35 (e.g., Camponotus (Myrmamblys), C. (Myrmentoma) and C. (Pseudocolobopsis)) clypeus tending to be relatively broad, such that CLW/CLL 1.25–1.62, although exceptions occur (e.g., in some C. (Pseudocolobopsis) species) (Figures 14–15); antennal insertions usually occurring in front of midlength of frontal carinae; clypeus typically with prominent anterolateral extensions (Figure 15) ... Camponotus 3 With conspicuous long setae, gracile legs, and a shield-shaped clypeus with prominent anterolateral extensions (Figure 16) ... Camponotus chloroticus - Without the combination of conspicuous long setae and gracile legs; clypeus lacking prominent anterolateral extensions (Figures 8–11) ... Colobopsis 4 Small species, HW 0.68–1.04; antennal insertions more widely separated (ASM/HW 0.34–0.39 and ASM/CLW 0.64–0.77) (Figures 18–19); clypeus tending to be less broad (CLW/CLL 1.15–1.40) ... Colobopsis - Small to medium-sized species, HW 0.75–2.10; antennal insertions less well separated (ASM/HW 0.25–0.29 and ASM/CLW 0.46–0.55); clypeus varying in shape, but if HW < 1.05 (e.g., Camponotus pulchellus complex) (Figure 17) then clypeus tending to be broader (CLW/CLL 1.25–1.60) ... Camponotus Fossils Fossils are known from: Aix-en-Provence, France (Late Oligocene), Baltic amber (Bartonian, Middle to Late Eocene), Barstow Formation, California, United States (Burdigalian to Langhian, Early to Middle Miocene), Bembridge Marls, Isle of Wight, UK (Priabonian, Late Eocene), Bitterfeld amber (Bartonian, Middle to Late Eocene), Brunn-Vösendorf, Austria (Late Miocene), Brunstatt, Haut-Rhin, France (Early Oligocene), Danish-Scandinavian amber (Bartonian, Middle to Late Eocene), Decín, Czech Republic (Early Miocene), Florissant, Colorado, United States (Late Eocene), Fushun amber, Liaoning, China (Ypresian, Early Eocene), Green River Formation, Colorado, United States (Lutetian, Middle Eocene), Joursac, Auvergne, France (Late Miocene), Montagne d'Andance, Saint-Bauzile, Ardèche, France (Early Turolian, Late Miocene), Oeningen, Switzerland (Messinian, Late Miocene), Radoboj, Croatia (Burdigalian, Early Miocene), Rovno amber (Priabonian, Late Eocene), Shanwang, China (Early Miocene). Biology Nesting Habit There is great diversity of nest architecture: in ground, in dead wood, or structures constructed outside tree trunks and branches using carton (rigid material composed of masticated and moistened plant fibres) or silk. Species belonging to three subgenera (Karavaievia in Asia; Dendromyrmex and Myrmobrachys in South America) use last-instar larvae to weave their nests, and incorporate sand particles, detritus and bits of plants into the silk sheet (e.g. Camponotus texens and Camponotus gombaki; Maschwitz et al. 1985). See also Camponotus nitidior, Camponotus chartifex, Camponotus senex A twig-nesting species from Cairns, Queensland, Australia. Ant Gardens Four unidentified species of Camponotus from peninsular Malaysia are known to form ant-gardens (i.e., they are able to initiate ant gardens or are restricted to ant gardens) (Kaufmann at al., 2001; Kaufmann, 2002 (noted as ant-garden initiator); Kaufmann & Maschwitz, 2006; Orivel & Leroy, 2011). Life History Traits
Nomenclature The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.
Common Names for Camponotus Subgenera A mnemonic list of common names for the subgenera of Camponotus may be found here. Subgenus Camponotus True Carpenter Ants Diagnosis of subgenus: Emery (1925) - "Worker and queen. - Large species; caste polymorphism and other characteristics as in the subgenus Tanaemyrmex, except for the following differences: Head of minors Alpha-form (see Tanaemyrmex for head shape classification), but head generally shorter and posterior head margin straighter; head of major worker much less enlarged and less emarginate posteriorly; head of queens similar to those of majors. Clypeus lacking medial carina nor usually with anterior lobe, although weakly-produced and more-or-less rectangular (C. japonicus, Mayr) or rounded (C. sansabeanus, Buckley) lobe may be present [note: C. sansabeanus is currently classified as Tanaemyrmex]; anterior margin of clypeus entire, lacking medial notch. Mandibles strongly arched, usually with 4 or 5 teeth, sometimes with 6. Mesosomal dorsum arched, always continuous in profile; pronotal dorsum rounded or sometimes depressed in majors, with slightly prominent humeri. Male. - Caracteres of Tanaemyrmex. Note that the subgenera Camponotus, Tanaemyrmex, Myrmosericus, and Myrmothrix constitute a group: they are separated by characters of little importance and often offer transitions from one group to another. Geographical distribution of species. - Holarctic region, except Mediterranean Africa; C. punctatissimus, Emery, is found in the north of Indomalaya." (Translated and edited by B. E. Boudinot, 15 February 2017.) Taxonomic History of Camponotus The taxonomic history of Camponotus may be divided into four phases, or eras, based on the kind of work published during these time periods. Phase I ("Archaic Era") 1861–1896 (Beginning: Mayr 1861; end: Emery 1896). The beginning of subgeneric work on Camponotus started with Mayr's (1861) key to the European ant fauna, wherein he recognized the cork-head ants, Colobopsis, as a distinct genus for the first time. Most of the work during this period was regional in nature, treating Europe, the Mediterranean, Madagascar, Russia, India, Sri Lanka, and North America. Only one explicit subgenus, Mayria was designated, while two taxa currently treated as subgenera were described: the American weaving ants, Dendromyrmex, and big-nose carpenter ants, Rhinomyrmex. In 1896, Emery proposed the first global, synoptic classification of Camponotus; previously no higher classification was in place, excepting recognition Mayria. In this work, all of the groups Emery distinguished were treated informally. Specifically, Emery recognized three principal groups (cohorts), which he divided into 26 subgroups which he termed manipoli. This work marked the beginning of the second phase of higher classification within Camponotus, wherein formal treatments of these groups were published. Phase II ("Classical Era") 1896–1928 (First classificatory publication of era for Camponotus: Emery, 1898; last: Santschi, 1928). In addition to the continuation of global exploration marked by regional works (South America, India, Sri Lanka, Myanmar, Palearctic [numerous], North America, Afrotropics [particularly South Africa], and the Malagasy region), this period is distinguished by greater focus on the subgenera, with Forel (1914), Emery (1920), Wheeler (1922), Santschi (1921), and Arnold (1922) expanding upon and refining Emery's system. In 1912, Forel formalized the Emery's (1896) higher classification by recognizing 16 subgenera, but without designating types. Wheeler (1913) sought to correct this by explicitly designating type species, but unfortunately this work was overlooked when Forel (1914) redundantly designated type specimens for his subgenera. This error of omission was replicated by Emery (1920), wherein he revised his initial classification, recognizing a total of 34 subgenera, as well as excluding Phasmomyrmex and species belonging to Notostigma (Melophorini), which he described as new. Wheeler (1921) put the nail in the coffin for the issue of confusion over subgeneric type species in a pointedly titled article, Professor Emery's subgenera of the genus Camponotus Mayr. The capstone to this second phase was Emery' (1925) final ,monumental contribution to Wytsmann's Genera Insectorum, which is still the most significant work on the internal classification of Camponotus to date. In this work, Emery synthesized all previous taxonomic works on Camponotus, particularly those of Forel and Wheeler, revised the delimitation of all subgenera, including subgroups, and provided keys to and diagnoses of all subgenera, all of which amounted to an effort which has not been matched. Moreover, Emery proposed his final phylogenetic hypothesis for the genus (see Emery's Phylogenetic Hypothesis below), which is distinguished from hypotheses he proposed for other groups by the strong biogeographic structure he postulated for the group. In 1926 and 1928, Santschi published the final two works formally altering Emery's (1925) system; both of these works were minor acts of splitting, and neither revised Emery's keys. Phase III ("Modern Era") 1928–2006 (Beginning: Santschi, 1928; end: Moreau et al. and Brady et al. 2006) After Emery (1925) and Santschi (1926, 1928), no further works were published which formally modified the subgeneric system of Camponotus until the fourth era of the internal classification of the genus (see Phase IV below). For almost 90 years, there has been, effectively, no major attempt to understand the diversity of Camponotus globally, with the major publications of the era for the genus being regional catalogs (Kempf 1972 Neotropics (followed by Brandao’s update in 1991), Taylor & Brown 1985 Australia (followed by Shattuck 1999), and Bolton’s 1995 global catalog). Virtually all of the taxonomic work on Camponotus during this time were regional, which in sum span the globe, but with global revisions restricted to minor groups of species (i.e., the groups or complexes of herculeanus, nigriceps, fulvopilosus, weiderkehri, and perjerus), as well as regionally-restricted and well-defined or easily-recognizable subgenera (e.g., Dendromyrmex, Karavaievia, Myrmentoma, Myrmostenus, Myrmothrix). In the late ‘90s and early 2000’s, Shattuck and McArthur treated a number Australian groups which they recognized morphologically but in which they disregarded the subgeneric classification. The end of this era is marked by the first global phylogenies of the Formicidae based on molecular data, published by Moreau et al. (2006) and Brady et al. (2006). Phase IV ("Molecular Era") 2006–present. The current era of classification, with respect to Camponotus, is nascent. Blaimer et al. (2015) published two phylogenies of the Formicinae using “traditional” Sanger sequencing and “next-gen” Illumina sequencing of ultraconserved elements (UCEs). This work was followed shortly by a formal reclassification of the subfamily by Ward et al. (2016), in which a number of taxonomic changes were made to relieve Camponotus and particular subgenera of paraphyly (i.e., Colobopsis and Dinomyrmex revived status as genera, Myrmogonia synonymized with Colobopsis, Phasmomyrmex made subgenus of Camponotus).Bill MacKay is still working on his revision of Camponotus of the Americas, and at present, no major phylogenetic work is underway for Camponotus, beyond refining the placement of species belonging to Colobopsis. (Section by B. E. Boudinot, 19 February 2017.) Emery's Phylogenetic Hypothesis for Camponotus In Emery's (1925) treatment of Camponotus in Genera Insectorum, he proposed a final, revised hypothesis of the internal relationships of the genus; an earlier version was published in Emery (1920). An important facet of this hypothesis, translated from p. 61, is Emery's "opinion that Camponotus underwent independent evolution in the Old and New Worlds". Emery postulated seven genus groups within Camponotus, with explicit morphologically-transitional lineages. These groups are indicated below: Group I: Myrmopiromis (Afrotropical and Malagasy regions, a few species from India) 1. Myrmopiromis Group II: Myrmosaulus (Malaysia, Australia, Madagascar)1. Myrmosaulus Group III: Myrmamblys Myrmopsamma subgroup (Africa) 1. Myrmopsamma Karavaievia subgroup (Indomalaya, etc.) 1. Karavaievia Mayria subgroup (Afrotropical, Malagasy) 1. Mayria Miscellaneous subgroup 1. Myrmentoma Subgroup transitional to Myrmaphaenus via Tanaemyrmex 1. Myrmamblys (Old World) [note: split into Myrmespera, Myrmopelta, Myrmotemnus, Paramyrmamblys, and Myrmamblys sensu stricto by Santschi (1926).] Group IV: Tanaemyrmex (Cosmopolitan) 1. Tanaemyrmex [note: Emery's species groups may be particularly useful for understanding, and dividing, this probably "plesiotypic" taxon] Group V: Myrmoxygenys group (Transitional between Myrmamblys and Myrmaphaenus groups via Tanaemyrmex) 1. Myrmoxygenys (Afrotropics) Group VI: Myrmaphaenus Subgroup transitional to Myrmamblys group via Tanaemyrmex 1. Camponotus (Holarctic) (sensu lato, including Nearctic Tanaemyrmex and caryae and lateralis groups of Myrmentoma, in part. Neotropical subgroup, transitional to Myrmobrachys group 1. Pseudocolobopsis Group VII: Myrmobrachys Subgroup transitional to Myrmaphaenus group 1. Myrmobrachys (Section by B. E. Boudinot, 19 February 2017.). |
Distribution and Richness based on AntMaps Synonyms Dolophra (Wu, J. & Wang, 1994)Myrmocamelus (Forel, 1914) Myrmolophus (Emery, 1920) Myrmoturba (Forel, 1912) Neocolobopsis (Borgmeier, 1928) Neomyrmamblys (Wheeler, W.M., 1921) Orthonotus (Ashmead, 1905) Paleosminthurus (Pierce & Gibron, 1962) Paracolobopsis (Emery, 1920) Shanwangella (Zhang, J., 1989) Subgenera Camponotus (Mayr, 1861) Diversity 1,501 species
Cvens Sesaxeb ჩვენი საიტი მოიცავს საქართველოს ფლორას და ფაუნაას. თქვენ მიიღებთ ბევრ საინტერესო ინფორმაციას ბუნებასა და დედამიწის შესახებ მათ შორის ცხოველთა სამყაროზე. ასევე გაეცნობით უიშვიათესი ჯიშის მცენარეებს. ჩვენი საიტის მეშვეობით თქვენ მიიღებთ ურიცხვ ინფორამციას ჩვენი ქვეყნის ბუნების შესახებ.
"წინამდებარე ტექსტი/ფოტოები გადმოკოპირებულია შემდეგი ინტერნეტ წყაროებიდან: https://apa.gov.ge/ge/, https://www.wikipedia.org/. აღნიშნულთან დაკავშირებით რაიმე სახის პრეტენზიის არსებობის შემთხვევაში, გთხოვთ დაუკავშირდეთ ადმინისტრაციას მითითებულ საკონტაქტო რეკვიზიტებზე" | ||||||||||||||||||||